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Many of the above groups are paraphyletic, in that they have given rise to successive groups: Agnathans are ancestral to Chondrichthyes, who again have given rise to Acanthodiians, the ancestors of Osteichthyes.
With the arrival of phylogenetic nomenclature, the fishes has been split up into a more detailed scheme, with the following major groups: † – indicates extinct taxon Some palaeontologists contend that because Conodonta are chordates, they are primitive fish.
Phylogenetic research in 19 supported the idea that the hagfish and the lampreys form a natural group, the Cyclostomata, that is a sister group of the Gnathostomata.
The various fish groups account for more than half of vertebrate species.
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